The animal kingdom is characterized by an incredible breadth of biological forms, ranging from microscopic organisms to massive multicellular structures. Among the Bilateria—animals possessing bilateral symmetry, three tissue layers, a coelom, and a complete gut—two major lineages emerge based on their embryonic development: the Protostomes and the Deuterostomes. Within the Protostome lineage, we find some of the most successful and diverse organisms on Earth, most notably the members of Phylum Arthropoda.
The Protostome Lineage and Ecdysozoans
To understand arthropods, we must first understand their place in the evolutionary tree. Protostomes are defined by several key developmental characteristics. During early embryonic development, the blastopore (the first opening in the embryo) becomes the mouth. Furthermore, protostomes undergo determinate cleavage (also known as mosaic cleavage), where the fate of each embryonic cell is determined very early in development. This means that if a cell is removed from a developing protostome embryo, the resulting organism will not develop normally.
Within the protostomes, there is a major group known as the Ecdysozoans. Unlike Lophotrochozoans, which grow incrementally, Ecdysozoans grow through a process called ecdysis. Ecdysis is the periodic molting of a hard exoskeleton or cuticle, which allows the animal to increase in size.
One prominent group within the Ecdysozoans is Phylum Nematoda, commonly known as "roundworms". Nematodes are pseudocoelomates, meaning they possess a false body cavity. They can be free-living or parasitic, causing various human health issues such as elephantiasis, hookworm infections, pinworms, and guinea worm disease.
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