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Sweet clover에 곰팡이 1) _______________________ 가 감염되면 식물의 Coumarin이 2) ________________로 전환된다. Coumarin 자체는 혈액응고 억제작용이 없으나 곰팡이에 의해 4-hydroxycoumarin 같은 화합물로 대사되고, 이어서 미생물이 생성한 formaldehyde와 반응에 2) _____________로 생합성된다.

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