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1-grain de pollen individualisés (monades) le plus souventRelief tectum peu marqué (poacées, arbres a chatons)Ornementation importante (la plupart des angioS)2-Grains de pollen groupés par 4 : tétrades Indissociables 3-Grain de pollen en groupes supérieurs a 4 : polyades Ex : alibizia (fabacées)4-Grain de pollen agglomérés en pollinies Ex : orchidacées, asclépiadacées
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